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    <title>UTas ePrints - Early Tertiary Macrofossils of Proteaceae from Tasmania</title>
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    <meta content="Carpenter, Raymond J." name="eprints.creators_name" />
<meta content="Jordan, Gregory J." name="eprints.creators_name" />
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<meta content="Proteaceous leaves from two Early Tertiary sites in Tasmania are described and discussed.  Based on gross leaf morphological and cuticular characters, 15 species are recognised from Oligocene lacustrine sediments at Cethana.  Specimens of two taxa are not distinguishable from the extant species Telopea truncata from Tasmania and Lomatia fraxinifolia from north-eastern Queensland, and are therefore assigned to these species.  Two new species of Banksieaephyllum are recognised.  Ten other taxa are difficult to identify to existing genera, and are therefore referred to a new genus Proteaciphyllum.  They all possess features typical of subfamily Grevilleoideae.  Other, less well preserved, but probable Proteaceae from Cethana are also described.  Specimens from the Leven River deposit, probably also of Oligocene age, are assigned to a new species of Orites, O. excelsoides.  This species is closely related to extant O. excelsa from rainforests of north-eastern New South Wales and north-eastern Queensland.  Cethana has by far the highest diversity of Proteaceae of any fossil floras described to date.  The fossils also demonstrate the past association of now geographically remote taxa, and the prevalence of sclero- and xeromorphy in the family by the Oligocene." name="eprints.abstract" />
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<meta content="Ancibor, E. (1989).  Determinacion xilologica de una raiz petrificada de Proteaceae de la Formacion Rio Turbio (Eoceno) Santa Cruz, Argentina.  Ameghiniana 25, 289-295.
Beadle, N. C. W. (1966).  Soil phosphate and its role in molding segments of the Australian flora and vegetation, with special reference to xeromorphy and sclerophylly.  Ecology 47, 992-1007.
Blackburn, D. T. (1981).  Tertiary megafossil flora of Maslin bay, South Australia:  numerical taxonomic study of selected leaves.  Alcheringa 5, 9-28.
Blackburn, D. T., and Sluiter, I. R. K. (1994).  The Oligo-Miocene coal floras of southeastern Australia.  In 'Australian Vegetation History: Cretaceous to Recent'.  (Ed. R. S. Hill.)  pp. 328-367.  (Cambridge University Press: Cambridge.)
Carpenter, R. J. (1991a).  'Palaeovegetation and Environment at Cethana, Tasmania.'  PhD Thesis, University of Tasmania, Hobart.
Carpenter, R. J. (1991b).  Macrozamia from the Early Tertiary of Tasmania and a study of the cuticles of extant species.  Australian Systematic Botany 4, 433-444.
Carpenter, R. J. (1994).  Cuticular morphology and aspects of the ecology and fossil history of North Queensland rainforest Proteaceae.  Botanical Journal of the  Linnean Society 116, 249-303.
Carpenter, R. J., and Buchanan, A. M. (1993).  Oligocene leaves, fruit and flowers of the Cunoniaceae from Cethana, Tasmania.  Australian Systematic Botany 6, 91-109.
Carpenter, R. J., and Hill, R. S. (1988).  Early Tertiary Lomatia (Proteaceae) macrofossils from Tasmania, Australia.  Review of Palaeobotany and  Palynology 56, 141-150.
Carpenter, R. J., and Pole, M. (1995).  Eocene plant fossils from the Lefroy and Cowan paleodrainages, Western Australia.  Australian Systematic Botany 8, 1107-1154.
Carpenter, R. J., Hill, R. S., and Jordan, G. J. (1994).  Cenozoic vegetation in Tasmania: macrofossil evidence.  In 'Australian Vegetation History: Cretaceous  to Recent'.  (Ed. R. S. Hill.)  pp. 276-298.  (Cambridge University Press:  Cambridge.)
Carpenter, R. J., Jordan, G. J., and Hill, R. S. (1994).  Banksieaephyllum taylorii  (Proteaceae) from the Late Paleocene of New South Wales and its relevance to the origin of Australia's scleromorphic flora.  Australian Systematic Botany 7, 385-392.
Christophel, D. C. (1984).  Early Tertiary Proteaceae: The first floral evidence for the Musgraveinae.  Australian Journal of Botany 32, 177-186.
Christophel, D. C. (1989).  Evolution of the Australian flora through the Tertiary.  Plant Systematics and Evolution 162, 63-78.
Christophel D. C, and Greenwood D. R. (1988).  A comparison of tropical rainforest and Tertiary fossil leaf beds.  Proceedings of the Ecological Society of Australia 15, 139-148.
Christophel D. C., Scriven L. J., and Greenwood D. R. (1992).  An Eocene megafossil flora from Nelly Creek, South Australia.  Transactions of the Royal Society of South Australia 116, 65-76.
Cookson, I. C., and Duigan, S. L. (1950).  Fossil Banksieae from Yallourn, Victoria, with notes on the morphology and anatomy of living species.  Australian Journal of Scientific Research, Series B 3, 133-165.
Dettmann, M. E. (1989).  Antarctica: Cretaceous cradle or grave of austral temperate  rainforests?  In 'Origins and Evolution of the Antarctic Biota'.  (Ed. J. A. Crame.)  Geological Society of London Special Publication 47, 89-105.
Dettmann, M. E., and Jarzen, D. M. (1991).  Pollen evidence for Late Cretaceous differentiation of Proteaceae in southern polar forests.  Canadian Journal of  Botany 69, 901-6.
Douglas, A. W. (1995).  Affinities.  Flora of Australia 16, 6-14.
Frenguelli, J. (1943).  Proteaceás del Cenozoico de Patagonia.  Notas del Museo de La Plata, Paleontologia 52, 201-213.
George, A. S., and Hyland, B. P. M. (1995).  Orites.  Flora of Australia 16, 346-352.
Hickey, L. J. (1979).  A revised classification of the architecture of dicotyledonous leaves.  In 'Anatomy of the Dicotyledons, Vol. 1'.  (Eds C. R. Metcalfe and L.  W. Chalk.)  pp. 25-39.  (Clarendon Press: Oxford.)
Hill, R. S. (1990).  Tertiary Proteaceae in Australia: A re-investigation of Banksia adunca and Dryandra urniformis.  Proceedings of the Royal Society of Victoria 102, 23-28.
Hill, R. S. (1994).  The history of selected Australian taxa.  In  'Australian Vegetation History: Cretaceous to Recent'.  (Ed. R. S. Hill.)  pp. 390-419.  (Cambridge  University Press: Cambridge.)
Hill, R. S., and Christophel, D. C. (1988).  Tertiary leaves of the tribe Banksieae (Proteaceae) from south-eastern Australia.  Botanical Journal of the Linnean Society 97, 205-227.
Hill, R. S., and Merrifield, H. E. (1993).  An Early Tertiary macroflora from West Dale, southwestern Australia.  Alcheringa 17, 285-326.
Hill, R. S., Scriven, L. J. and Jordan, G. J. (1995).  The fossil record of Australian Proteaceae.  Flora of Australia 16, 21-30.
Johnson, L. A. S., and Briggs, B. G. (1975).  On the Proteaceae, the evolution and classification of a southern family. Botanical Journal of the Linnean Society 70, 83-182.
Johnson, L. A. S., and Briggs, B. G. (1981).  Three old southern families - Myrtaceae, Proteaceae and Restionaceae. In 'Ecological Biogeography of Australia'.  (Ed. A. Keast.)  pp. 429-469.  (W. Junk: The Hague.)
Jordan, G. J. (1995).  Early-Middle Pleistocene leaves of extinct and extant Proteaceae from western Tasmania, Australia.  Botanical Journal of the Linnean Society 118, 19-35.
Jordan, G. J., Carpenter, R. J., and Hill, R. S. (1991).  Late Pleistocene vegetation and climate near Melaleuca Inlet, south-western Tasmania.  Australian Journal of  Botany 39, 499-511.
Lange, R. T. (1978).  Some Eocene leaf fragments comparable to Proteaceae.  Journal of the Royal Society of Western Australia 60, 107-114.
Macphail, M. K., Colhoun, E. A., Kiernan, K., and Hannan, D. (1993).  Glacial climates in the Antarctic region during the late Paleogene: Evidence from northwest Tasmania, Australia.  Geology 21, 145-148.
Macphail, M. K., Alley, N. F., Truswell, E. M., and Sluiter, I. R. K. (1994).  Early Tertiary vegetation: evidence from spores and pollen. In 'Australian Vegetation History: Cretaceous to Recent'.  (Ed. R. S. Hill.)  pp. 189-261.  (Cambridge University Press: Cambridge.)
Martin, H. A. (1982).  Changing Cenozoic barriers and the Australian paleobotanical record.  Annals of the Missouri Botanical Garden 69, 625-667.
Pole, M. (1992).  Eocene vegetation from Hasties, north-eastern Tasmania.  Australian Systematic Botany 5, 431-475.
Specht, R. L. (1988).  Origin and evolution of terrestrial plant communities in the wet-dry tropics of Australia.  Proceedings of the Ecological Society of Australia 15, 19-30.
Turner, I. M. (1994).  Sclerophylly: primarily protective?  Functional Ecology 8, 669-675.
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<meta content="Proteaceous leaves from two Early Tertiary sites in Tasmania are described and discussed.  Based on gross leaf morphological and cuticular characters, 15 species are recognised from Oligocene lacustrine sediments at Cethana.  Specimens of two taxa are not distinguishable from the extant species Telopea truncata from Tasmania and Lomatia fraxinifolia from north-eastern Queensland, and are therefore assigned to these species.  Two new species of Banksieaephyllum are recognised.  Ten other taxa are difficult to identify to existing genera, and are therefore referred to a new genus Proteaciphyllum.  They all possess features typical of subfamily Grevilleoideae.  Other, less well preserved, but probable Proteaceae from Cethana are also described.  Specimens from the Leven River deposit, probably also of Oligocene age, are assigned to a new species of Orites, O. excelsoides.  This species is closely related to extant O. excelsa from rainforests of north-eastern New South Wales and north-eastern Queensland.  Cethana has by far the highest diversity of Proteaceae of any fossil floras described to date.  The fossils also demonstrate the past association of now geographically remote taxa, and the prevalence of sclero- and xeromorphy in the family by the Oligocene." name="DC.description" />
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    <h1 class="ep_tm_pagetitle">Early Tertiary Macrofossils of Proteaceae from Tasmania</h1>
    <p style="margin-bottom: 1em" class="not_ep_block"><span class="person_name">Carpenter, Raymond J.</span> and <span class="person_name">Jordan, Gregory J.</span> (1997) <xhtml:em>Early Tertiary Macrofossils of Proteaceae from Tasmania.</xhtml:em> Australian Systematic Botany, 10 (4). pp. 533-563.</p><p style="margin-bottom: 1em" class="not_ep_block"></p><table style="margin-bottom: 1em" class="not_ep_block"><tr><td valign="top" style="text-align:center"><a href="http://eprints.utas.edu.au/1729/1/Cethana_Proteaceae.pdf"><img alt="[img]" src="http://eprints.utas.edu.au/style/images/fileicons/application_pdf.png" border="0" class="ep_doc_icon" /></a></td><td valign="top"><a href="http://eprints.utas.edu.au/1729/1/Cethana_Proteaceae.pdf"><span class="ep_document_citation">PDF</span></a> - Full text restricted - Requires a PDF viewer<br />820Kb</td><td><form method="get" accept-charset="utf-8" action="http://eprints.utas.edu.au/cgi/request_doc"><input value="2225" name="docid" accept-charset="utf-8" type="hidden" /><div class=""><input value="Request a copy" name="_action_null" class="ep_form_action_button" onclick="return EPJS_button_pushed( '_action_null' )" type="submit" /> </div></form></td></tr></table><p style="margin-bottom: 1em" class="not_ep_block">Official URL: <a href="http://dx.doi.org/10.1071/SB96016">http://dx.doi.org/10.1071/SB96016</a></p><div class="not_ep_block"><h2>Abstract</h2><p style="padding-bottom: 16px; text-align: left; margin: 1em auto 0em auto">Proteaceous leaves from two Early Tertiary sites in Tasmania are described and discussed.  Based on gross leaf morphological and cuticular characters, 15 species are recognised from Oligocene lacustrine sediments at Cethana.  Specimens of two taxa are not distinguishable from the extant species Telopea truncata from Tasmania and Lomatia fraxinifolia from north-eastern Queensland, and are therefore assigned to these species.  Two new species of Banksieaephyllum are recognised.  Ten other taxa are difficult to identify to existing genera, and are therefore referred to a new genus Proteaciphyllum.  They all possess features typical of subfamily Grevilleoideae.  Other, less well preserved, but probable Proteaceae from Cethana are also described.  Specimens from the Leven River deposit, probably also of Oligocene age, are assigned to a new species of Orites, O. excelsoides.  This species is closely related to extant O. excelsa from rainforests of north-eastern New South Wales and north-eastern Queensland.  Cethana has by far the highest diversity of Proteaceae of any fossil floras described to date.  The fossils also demonstrate the past association of now geographically remote taxa, and the prevalence of sclero- and xeromorphy in the family by the Oligocene.</p></div><table style="margin-bottom: 1em" border="0" cellpadding="3" class="not_ep_block"><tr><th valign="top" class="ep_row">Item Type:</th><td valign="top" class="ep_row">Article</td></tr><tr><th valign="top" class="ep_row">Additional Information:</th><td valign="top" class="ep_row">The definitive version is available online at http://www.publish.csiro.au/nid/150.htm</td></tr><tr><th valign="top" class="ep_row">Keywords:</th><td valign="top" class="ep_row">biological diversity, scleromorphy, sclerophylly, fossil</td></tr><tr><th valign="top" class="ep_row">Subjects:</th><td valign="top" class="ep_row"><a href="http://eprints.utas.edu.au/view/subjects/270705.html">270000 Biological Sciences &gt; 270700 Ecology and Evolution &gt; 270705 Palaeoecology</a><br /><a href="http://eprints.utas.edu.au/view/subjects/260112.html">260000 Earth Sciences &gt; 260100 Geology &gt; 260112 Palaeontology</a><br /><a href="http://eprints.utas.edu.au/view/subjects/270401.html">270000 Biological Sciences &gt; 270400 Botany &gt; 270401 Plant Systematics, Taxonomy and Phylogeny</a></td></tr><tr><th valign="top" class="ep_row">Collections:</th><td valign="top" class="ep_row">UNSPECIFIED</td></tr><tr><th valign="top" class="ep_row">ID Code:</th><td valign="top" class="ep_row">1729</td></tr><tr><th valign="top" class="ep_row">Deposited By:</th><td valign="top" class="ep_row"><span class="ep_name_citation"><span class="person_name">dr gregory j jordan</span></span></td></tr><tr><th valign="top" class="ep_row">Deposited On:</th><td valign="top" class="ep_row">02 Sep 2007</td></tr><tr><th valign="top" class="ep_row">Last Modified:</th><td valign="top" class="ep_row">11 Feb 2008 11:04</td></tr><tr><th valign="top" class="ep_row">ePrint Statistics:</th><td valign="top" class="ep_row"><a target="ePrintStats" href="/es/index.php?action=show_detail_eprint;id=1729;">View statistics for this ePrint</a></td></tr></table><p align="right">Repository Staff Only: <a href="http://eprints.utas.edu.au/cgi/users/home?screen=EPrint::View&amp;eprintid=1729">item control page</a></p>
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